The rigid shell means turtles cannot breathe as other reptiles do, by changing the volume of their chest cavities via expansion and contraction of the ribs. Instead, they breathe in two ways. First, they employ buccal pumping, pulling air into their mouths, then pushing it into their lungs via oscillations of the floor of the throat. Secondly, when the abdominal muscles that cover the posterior opening of the shell contract, the internal volume of the shell increases, drawing air into the lungs, allowing these muscles to function in much the same way as the mammalian diaphragm.
The shape of the shell gives helpful clues about how a turtle lives. Most tortoises have a large, dome-shaped shell that makes it difficult for predators to crush the shell between their jaws. One of the few exceptions is the African pancake tortoise, which has a flat, flexible shell that allows it to hide in rock crevices. Most aquatic turtles have flat, streamlined shells which aid in swimming and diving. American snapping turtles and musk turtles have small, cross-shaped plastrons that give them more efficient leg movement for walking along the bottom of ponds and streams.
The color of a turtle's shell may vary.